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Several options exist for deriving management reference points and catch advice in SPoRC. In this vignette, we will first discuss the mathematical details for deriving different management reference points, and then demonstrate how catch advice might be developed with estimated reference points.

Reference Points

Deriving reference points in SPoRC is generally divided into single-region and multi-region reference points, along with spawning potential ratio (SPRSPR) and maximum sustainable yield (MSYMSY) reference points. In the following, we will discuss the mathematical details pertaining to these key management quantities.

Single Region

Single-region reference points can currently be derived for SPRSPR reference points, which estimate the fishing mortality rate that reduces spawning biomass per recruit (SSBPRSSBPR) to x%x\%\ of the unfished level. Additionally, MSYMSY reference points based on a Beverton-Holt relationship, which estimate the fishing mortality rate that maximizes long-term yield, can also be derived if density-dependence is assumed.

Spawning Potential Ratio

To derive SPRSPR reference points, a target percentage must be specified, representing the SSBPRSSBPR under fishing relative to the unfished level. Following that, several additional quantities are needed to compute these reference points. These include:

  • the relative fishing mortality between fishery fleets (Fratf=Fmorty=ny,f/fFmorty=ny,fFrat_{f} = Fmort_{y = n_{y},f}/\sum_{f}^{}{Fmort_{y = n_{y},f}}),
  • fleet-specific fishery selectivity (Sely=y*a,s,fFsh\text{Sel}_{y = y^{*}a,s,f}^{\text{Fsh}}),
  • natural mortality for females (Natmorty=y*,a,s=1\text{Natmort}_{y = y^{*},a,s = 1}),
  • spawning weight at age for females (Wy=y*,a,s=1spawnW_{y = y^{*},a,s = 1}^{spawn}),
  • maturity at age for females (Maty=y*,a,s=1Mat_{y = y^{*},a,s = 1}),
  • spawn timing (tspwnt^{spwn}),
  • estimated recruitment values (Recy,s{Rec}_{y,s}),
  • the female recruitment sex ratio (ψy=y*,s=1\psi_{y = y^{*},s = 1}),
  • recruitment age (RecAgeRecAge), and
  • the first year in the recruitment vector (RecYearRecYear) to be used for mean recruitment calculations.

For relative fishing mortality between fleets, the terminal year is utilized for these calculations. In contrast, estimates based on a user-defined period (y*y^{*}; e.g., three-year average) are used for natural mortality, weight at age, maturity at age, and fleet-specific selectivity.

SPRSPR\ reference points are then derived by initially setting numbers per-recruit at the female recruitment sex-ratio:

Na=1fished=ψy=y*,s=1N_{a = 1}^{fished} = \psi_{y = y^{*},s = 1}

Na=1unfished=ψy=y*,s=1N_{a = 1}^{unfished} = \psi_{y = y^{*},s = 1}

where Na=1fishedN_{a = 1}^{fished} and Na=1unfishedN_{a = 1}^{unfished} are the fished and unfished numbers at age per-recruit, respectively. Subsequent numbers at age per-recruit are then computed with an exponential mortality model, where fished numbers at age per-recruit are decremented with both natural and fishing mortality, while unfished numbers at age per-recruit are only decremented with natural mortality. For NafishedN_{a}^{fished}, this is computed as:

Nafished={Na1fishedexp[(NatMorty=y*,a1,s=1+fFratfFxSely=y*,a1,s=1,fFsh)]if 2<a<a+Na1fishedexp((NatMorty=y*,a+1,s=1+fFratfFxSely=y*,a+1,s=1,fFsh))1exp((NatMorty=y*,a+,s=1+fFratfFxSely=y*,a+,s=1,fFsh))if a=a+N_{a}^{\text{fished}} = \left\{ \begin{matrix} N_{a - 1}^{\text{fished}} \cdot \exp\left\lbrack - \left( NatMort_{y = y^{*},a - 1,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{x} \cdot \text{Sel}_{y = y^{*},a - 1,s = 1,f}^{\text{Fsh}} \right) \right\rbrack & \text{if }2 < a < a_{+} \\ & \\ N_{a - 1}^{\text{fished}} \cdot \frac{\exp\left( - \left( NatMort_{y = y^{*},a_{+} - 1,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{x} \cdot \text{Sel}_{y = y^{*},a_{+} - 1,s = 1,f}^{\text{Fsh}} \right) \right)}{1 - exp\left( - \left( NatMort_{y = y^{*},a_{+},s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{x} \cdot \text{Sel}_{y = y^{*},a_{+},s = 1,f}^{\text{Fsh}} \right) \right)} & \text{if }a = a_{+} \\ \end{matrix} \right.\

where FxF_{x} represents the estimated fishing mortality rate that reduces the SSBPRSSBPR to x%x\%, relative to the unfished level. For Na=1unfishedN_{a = 1}^{unfished}, this is defined as:

Naunfished={Na1unfishedexp(NatMorty=y*,a1,s=1)if 2<a<a+Na1unfishedexp(NatMorty=y*,a+1,s=1)1exp(NatMorty=y*,a+,s=1)if a=a+N_{a}^{\text{unfished}} = \left\{ \begin{matrix} N_{a - 1}^{\text{unfished}} \cdot exp\left( - NatMort_{y = y^{*},a - 1,s = 1} \right) & \text{if }2 < a < a_{+} \\ & \\ N_{a - 1}^{\text{unfished}} \cdot \frac{\exp\left( - NatMort_{y = y^{*},a_{+} - 1,s = 1} \right)}{1 - exp\left( - NatMort_{y = y^{*},a_{+},s = 1} \right)} & \text{if }a = a_{+} \\ \end{matrix} \right.\

NafishedN_{a}^{\text{fished}} can then be converted to fished SSBPRSSBPR (SSBPRfished)SS{BPR}^{\text{fished}})\ with the following equation:

SSBPRfished=aNafishedWy=y*,a,s=1spawnMaty=y*,a,s=1exp(tspawnNatMorty=y*,a,s=1+fFratfFxSely=y*a,s=1,fFsh)SS{BPR}^{\text{fished}} = \sum_{a}^{}N_{a}^{\text{fished}} \cdot W_{y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{y = y^{*},a,s = 1} \cdot exp\left( - t^{spawn} \cdot NatMort_{y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{x} \cdot \text{Sel}_{y = y^{*}a,s = 1,f}^{\text{Fsh}} \right)

Similarly, Na=1unfishedN_{a = 1}^{unfished} can be converted to unfished SSBPRSSBPR (SSBPRunfished)SS{BPR}^{\text{unfished}}):

SSBPRunfished=aNaunfishedWy=y*,a,s=1spawnMaty=y*,a,s=1exp(tspwnNatMorty=y*,a,s=1)SS{BPR}^{\text{unfished}} = \sum_{a}^{}N_{a}^{\text{unfished}} \cdot W_{y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{y = y^{*},a,s = 1} \cdot exp\left( - t_{\text{spwn}} \cdot NatMort_{y = y^{*},a,s = 1} \right)

The SPRSPR rate can then be defined as the fraction of SSBPRfishedSS{BPR}^{\text{fished}} and SSBPRunfishedSS{BPR}^{\text{unfished}}:

SPR=SSBPRfishedSSBPRunfishedSPR = \frac{SS{BPR}^{\text{fished}}}{SS{BPR}^{\text{unfished}}}

FxF_{x} can then be solved for using a non-linear function minimizer by minimizing the following criteria:

Fx=argminF{100(SPR(F)x%)2}F_{x} = arg\min_{F}\left\{ 100 \cdot \left( \text{SPR}(F) - \text{x%} \right)^{2} \right\}

Biological SPRSPR-based reference points (BxB_{x}) can then be computed as:

Bx=SSBPRfishedRecYearnyRecAgeRecy,snyRecAgeRecYearB_{x} = SS{BPR}^{\text{fished}} \cdot \frac{\sum_{RecYear}^{n_{y} - RecAge}{Rec}_{y,s}}{n_{y} - RecAge - RecYear}

where SSBPRfishedSS{BPR}^{\text{fished}}\ is multiplied by mean recruitment over a user-defined period (i.e., nyRecAgeRecYearn_{y} - RecAge - RecYear).

Maximum Sustainable Yield (Beverton-Holt)

Deriving MSYMSY based reference points using a Beverton-Holt stock recruitment relationship involves maximizing the equilibrium yield per recruit (YPRYPR) and requires several additional inputs. These inputs include:

  • the relative fishing mortality between fishery fleets (Fratf=Fmorty=ny,f/fFmorty=ny,fFrat_{f} = Fmort_{y = n_{y},f}/\sum_{f}^{}{Fmort_{y = n_{y},f}}),
  • fleet-specific fishery selectivity (Sely=y*a,s,fFsh\text{Sel}_{y = y^{*}a,s,f}^{\text{Fsh}}),
  • natural mortality for females (Natmorty=y*,a,s=1\text{Natmort}_{y = y^{*},a,s = 1}),
  • spawning weight at age for females (Wy=y*,a,s=1spawnW_{y = y^{*},a,s = 1}^{spawn}),
  • maturity at age for females (Maty=y*,a,s=1Mat_{y = y^{*},a,s = 1}),
  • spawn timing (tspawn)t^{spawn}),
  • estimated recruitment values (Recy,s{Rec}_{y,s}),
  • the female recruitment sex ratio (ψy=y*,s=1\psi_{y = y^{*},s = 1}),
  • an estimate of virgin recruitment (μRec\mu^{Rec}), and
  • an estimate of steepness (hh).

MSYMSY reference points can then be derived using the standard per-recruit calculations, where the initial number of fished and unfished individuals are set at the female recruitment sex ratio:

Na=1fished=ψy=y*,s=1N_{a = 1}^{fished} = \psi_{y = y^{*},s = 1}

Na=1unfished=ψy=y*,s=1N_{a = 1}^{unfished} = \psi_{y = y^{*},s = 1}

The numbers-at-age can then be decremented following an exponential mortality model. For Na=1fishedN_{a = 1}^{fished}, this is given by:

Nafished={Na1fishedexp[(NatMorty=y*,a1,s=1+fFratfFmsySely=y*,a1,s=1,fFsh)]if 2<a<a+Na1fishedexp((NatMorty=y*,a+1,s=1+fFratfFmsySely=y*,a+1,s=1,fFsh))1exp((NatMorty=y*,a+,s=1+fFratfFmsySely=y*,a+,s=1,fFsh))if a=a+N_{a}^{\text{fished}} = \left\{ \begin{matrix} N_{a - 1}^{\text{fished}} \cdot \exp\left\lbrack - \left( NatMort_{y = y^{*},a - 1,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{msy} \cdot \text{Sel}_{y = y^{*},a - 1,s = 1,f}^{\text{Fsh}} \right) \right\rbrack & \text{if }2 < a < a_{+} \\ & \\ N_{a - 1}^{\text{fished}} \cdot \frac{\exp\left( - \left( NatMort_{y = y^{*},a_{+} - 1,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{msy} \cdot \text{Sel}_{y = y^{*},a_{+} - 1,s = 1,f}^{\text{Fsh}} \right) \right)}{1 - exp\left( - \left( NatMort_{y = y^{*},a_{+},s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{msy} \cdot \text{Sel}_{y = y^{*},a_{+},s = 1,f}^{\text{Fsh}} \right) \right)} & \text{if }a = a_{+} \\ \end{matrix} \right.\

where FmsyF_{msy} here represents the fishing mortality that would result in equilibrium yield being maximized. For Na=1unfishedN_{a = 1}^{unfished}, this is defined as:

Naunfished={Na1unfishedexp(NatMorty=y*,a1,s=1)if 2<a<a+Na1unfishedexp(NatMorty=y*,a+1,s=1)1exp(NatMorty=y*,a+,s=1)if a=a+N_{a}^{\text{unfished}} = \left\{ \begin{matrix} N_{a - 1}^{\text{unfished}} \cdot exp\left( - NatMort_{y = y^{*},a - 1,s = 1} \right) & \text{if }2 < a < a_{+} \\ & \\ N_{a - 1}^{\text{unfished}} \cdot \frac{\exp\left( - NatMort_{y = y^{*},a_{+} - 1,s = 1} \right)}{1 - exp\left( - NatMort_{y = y^{*},a_{+},s = 1} \right)} & \text{if }a = a_{+} \\ \end{matrix} \right.\

YPRYPR is then computed using Baranov’s catch equation:

Ca=fFratfFmsySely=y*,a,s=1,fFsh(NatMorty=y*,a,s=1+fFratfFmsySely=y*,a,s=1,fFsh)Nafished(1exp[(NatMorty=y*,a,s=1+fFratfFmsySely=y*,a,s=1,fFsh)])C_{a} = \frac{\sum_{f}^{}\text{Frat}_{f} \cdot F_{\text{msy}} \cdot \text{Sel}_{y = y^{*},a,s = 1,f}^{\text{Fsh}}}{\left( NatMort_{y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{\text{msy}} \cdot \text{Sel}_{y = y^{*},a,s = 1,f}^{\text{Fsh}} \right)} \cdot N_{a}^{\text{fished}} \cdot \left( 1 - exp\left\lbrack - \left( NatMort_{y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{\text{msy}} \cdot \text{Sel}_{y = y^{*},a,s = 1,f}^{\text{Fsh}} \right) \right\rbrack \right)

where CaC_{a} is the catch-at-age per-recruit resulting from Baranov’s catch equation. Equilibrium recruitment can then computed using the Beverton-Holt stock recruitment relationship. Estimates of equilibrium recruitment are needed to convert per-recruit quantities to yield and BmsyB_{msy} (the biomass that would achieve MSYMSY). Here, we first SSBPRfishedSS{BPR}^{\text{fished}} and SSBPRunfishedSS{BPR}^{\text{unfished}}, which are necessary quantities in the Beverton-Holt stock recruitment relationship:

SSBPRfished=aNafishedWy=y*,a,s=1spawnMaty=y*,a,s=1exp(tspawnNatMorty=y*,a,s=1+fFratfFmsySely=y*a,s=1,fFsh)SS{BPR}^{\text{fished}} = \sum_{a}^{}N_{a}^{\text{fished}} \cdot W_{y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{y = y^{*},a,s = 1} \cdot exp\left( - t^{spawn} \cdot NatMort_{y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{f} \cdot F_{msy} \cdot \text{Sel}_{y = y^{*}a,s = 1,f}^{\text{Fsh}} \right)

SSBPRunfished=aNaunfishedWy=y*,a,s=1spawnMaty=y*,a,s=1exp(tspwnNatMorty=y*,a,s=1)SS{BPR}^{\text{unfished}} = \sum_{a}^{}N_{a}^{\text{unfished}} \cdot W_{y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{y = y^{*},a,s = 1} \cdot exp\left( - t_{\text{spwn}} \cdot NatMort_{y = y^{*},a,s = 1} \right)

Equilibrium recruitment (EqRecEqRec) can then be derived with the following equation:

EqRec=4hμRecSSBPRfishedSSBPRunfished(1h)+SSBPRfished(5h1)EqRec = \frac{4 \cdot h \cdot \mu^{Rec} \cdot SS{BPR}^{\text{fished}}}{SS{BPR}^{\text{unfished}} \cdot (1 - h) + SS{BPR}^{\text{fished}} \cdot (5 \cdot h - 1)}

Yield and BmsyB_{msy} is then derived by multiplying the YPRYPR\ and SSBPRfishedSS{BPR}^{\text{fished}} by an estimate of equilibrium recruitment:

Yield=aCaWy=y*,a,s=1spawnEqRecYield = \sum_{a}^{}C_{a} \cdot W_{y = y^{*},a,s = 1}^{spawn} \cdot EqRec

Bmsy=SSBPRfishedEqRecB_{msy} = SS{BPR}^{\text{fished}} \cdot EqRec

Lastly, FmsyF_{msy} is solved for using a non-linear function minimizer by minimizing the following criteria (maximizing yield):

Fmsy=argminFYieldF_{msy} = arg\min_{F}\ Yield

Multi-Region

In general, multi region reference points can be computed in a similar manner as single region reference points. The additional complication when calculating spatial reference points includes the additional region subscript for all quantities, as well as the potential need to account for movement processes.

Spawning Potential Ratio
Independent

In the case where independent spatial regions are assumed (i.e., no movement occurs among regions), SPRSPR rates can be calculated independently for each region, which results in region-specific Fr,xF_{r,x} and Br,xB_{r,x} estimates. All calculations are derived in the same manner as computing SPRSPR rates in the single region case, with the exception that an additional subscript is added to all demographic rates. Following that, Fr,xF_{r,x} can then be solved for by minimizing the following criteria for each region:

Fr,x=argminFr{100(SPR(Fr)x%)2}F_{r,x} = arg\min_{F_{r}}\left\{ 100 \cdot \left( \text{SPR}\left( F_{r} \right) - x\%\ \right)^{2} \right\}

Regional biological SPRSPR-based reference points (BxB_{x}) can then be derived by multiplying SSBPRrfishedSS{BPR}_{r}^{\text{fished}} by regional mean recruitment over a user-defined period and the recruitment sex-ratio:

Br,x=SSBPRrfishedRecYearnyRecAgeRecr,y,snyRecAgeRecYearB_{r,x} = SS{BPR}_{r}^{\text{fished}} \cdot \frac{\sum_{RecYear}^{n_{y} - RecAge}{Rec}_{r,y,s}}{n_{y} - RecAge - RecYear}

Global

In contrast to computing reference points when assuming independent spatial dynamics, SPRSPR rates can also be computed globally, where movement occurs among regions. Given the assumption of global SPRSPR rates, this results in a global FxF_{x} estimate, but regional estimates of Br,xB_{r,x} because mean recruitment estimates are defined on a regional scale. Thus, the global SPRSPR solution results in a FxF_{x} that reduces the global SSBPRSSBPR to x%x\% of its unfished value, such that the aggregate spawning biomass reaches equilibrium at rBr,x\sum_{r}^{}B_{r,x} if applied over the long-term. However, this solution does not result in each region equilibrating at Br,xB_{r,x}. Note that the same value of FxF_{x} is applied to each region in the global SPRSPR case.

Deriving global SPRSPR reference points requires a different set of inputs. These include:

  • the relative fishing mortality between fishery fleets (Fratr,f=Fmortr,y=ny,f/fFmortr,y=ny,fFrat_{r,f} = Fmort_{r,y = n_{y},f}/\sum_{f}^{}{Fmort_{r,y = n_{y},f}}),
  • fleet-specific fishery selectivity (Selr,y=y*a,s,fFsh\text{Sel}_{r,y = y^{*}a,s,f}^{\text{Fsh}}),
  • natural mortality for females (Natmortr,y=y*,a,s=1\text{Natmort}_{r,y = y^{*},a,s = 1}),
  • spawning weight at age for females (Wr,y=y*,a,s=1spawnW_{r,y = y^{*},a,s = 1}^{spawn}),
  • maturity at age for females (Matr,y=y*,a,s=1Mat_{r,y = y^{*},a,s = 1}),
  • a matrix of movement values (𝐌y=y*,a,s=1\mathbf{M}_{y = y^{*},\, a,\, s = 1}),
  • spawn timing (tspwnt^{spwn}),
  • estimated recruitment values (Recr,y,s{Rec}_{r,y,s}),
  • the female recruitment sex ratio (ψr,y=y*,s=1\psi_{r,y = y^{*},s = 1}),
  • the recruitment proportions (apportionment) by area (ζr\zeta_{r}),
  • recruitment age (RecAgeRecAge), and
  • the first year in the recruitment vector (RecYearRecYear) to be used for mean recruitment calculations.

Global SPRSPR reference points are calculated by first setting the regional numbers-at-age per-recruit model to the estimated recruitment apportionment parameters (ζr\zeta_{r}) multiplied by the recruitment sex ratio, which sum to 1:

Nr,a=1fished=ζrψr,y=y*,s=1N_{r,a = 1}^{\text{fished}} = \zeta_{r} \cdot \psi_{r,y = y^{*},s = 1}

Nr,a=1unfished=ζrψr,y=y*,s=1N_{r,a = 1}^{\text{unfished}} = \zeta_{r} \cdot \psi_{r,y = y^{*},s = 1}

where Nr,a=1fishedN_{r,a = 1}^{\text{fished}} and Nr,a=1unfishedN_{r,a = 1}^{\text{unfished}} are the regional numbers-at-age per-recruit under fished and unfished conditions, respectively. Following the initialization of these quantities, movement dynamics are then applied:

𝐍afished=(𝐍afished)𝐌y=y*,a,s=1for a=amin,,A,amin={1if recruits move2otherwise\mathbf{N}_{a}^{\text{fished}} = \left( \mathbf{N}_{a}^{\text{fished}} \right)^{\top}\mathbf{M}_{y = y^{*},a,s = 1}\quad\text{for }a = a_{\min},\ldots,A,\quad a_{\min} = \left\{ \begin{matrix} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \\ \end{matrix} \right.\

𝐍aunfished=(𝐍aunfished)𝐌y=y*,a,s=1for a=amin,,A,amin={1if recruits move2otherwise\mathbf{N}_{a}^{\text{unfished}} = \left( \mathbf{N}_{a}^{\text{unfished}} \right)^{\top}\mathbf{M}_{y = y^{*},a,s = 1}\quad\text{for }a = a_{\min},\ldots,A,\quad a_{\min} = \left\{ \begin{matrix} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \\ \end{matrix} \right.\

In the above expressions, AA denotes the number of modeled ages (a+a_{+}) multiplied by 10. For example, if 30 ages are modeled, then A=300A = 300. This extension enables iterative propagation of the plus group under movement. For aa+a \geq a_{+}, all demographic and fishery quantities are held constant at the values for the last modeled age (a+a_{+}). Following the application of movement, an exponential mortality model is then applied. Nr,afishedN_{r,a}^{fished} is expressed as:

Nr,afished=Nr,a1fishedexp[(NatMortr,y=y*,a1,s=1+fFratr,fFxSelr,y=y*,a1,s=1,fFsh)]if 2<a<AN_{r,a}^{\text{fished}} = {N_{r,a - 1}^{fished} \cdot exp}\begin{matrix} \left\lbrack - \left( NatMort_{r,y = y^{*},a - 1,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{x} \cdot \text{Sel}_{r,y = y^{*},a - 1,s = 1,f}^{\text{Fsh}} \right) \right\rbrack \\ \text{if }2 < a < A \\ \end{matrix}

For Nr,aunfishedN_{r,a}^{\text{unfished}}, this is given by:

Nr,aunfished=Nr,a1unfishedexp[(NatMortr,y=y*,a1,s=1)],if 2<a<AN_{r,a}^{\text{un}\text{fished}} = {N_{r,a - 1}^{unfished} \cdot exp}\begin{matrix} \left\lbrack - \left( NatMort_{r,y = y^{*},a - 1,s = 1} \right) \right\rbrack,\ \ \ \text{if }2 < a < A \\ \end{matrix}

Regional fished and unfished numbers-at-age per-recruit can then be converted to SSBPR quantities. For regional fished SSBPR (SSBPRrfished)SS{BPR}_{r}^{\text{fished}}), this is written as:

SSBPRrfished=aNr,afishedWr,y=y*,a,s=1spawnMatr,y=y*,a,s=1exp(tspawnNatMortr,y=y*,a,s=1+fFratr,fFxSelr,y=y*a,s=1,fFsh)SS{BPR}_{r}^{\text{fished}} = \sum_{a}^{}N_{r,a}^{\text{fished}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{r,y = y^{*},a,s = 1} \cdot exp\left( - t^{spawn} \cdot NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{x} \cdot \text{Sel}_{r,y = y^{*}a,s = 1,f}^{\text{Fsh}} \right)

Regional unfished SSBPRSSBPR (SSBPRrunfished)SS{BPR}_{r}^{\text{unfished}}) is computed in a similar manner:

SSBPRrunfished=aNr,aunfishedWr,y=y*,a,s=1spawnMatr,y=y*,a,s=1exp(tspwnNatMortr,y=y*,a,s=1)SS{BPR}_{r}^{\text{unfished}} = \sum_{a}^{}N_{r,a}^{\text{unfished}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{r,y = y^{*},a,s = 1} \cdot exp\left( - t_{\text{spwn}} \cdot NatMort_{r,y = y^{*},a,s = 1} \right)

The global SPRSPR rate is then defined as:

SPR=rSSBPRrfishedrSSBPRrunfishedSPR = \frac{\sum_{r}^{}{SS{BPR}_{r}^{\text{fished}}}}{\sum_{r}^{}{SS{BPR}_{r}^{\text{unfished}}}}

FxF_{x} can then be solved for using a non-linear function minimizer by minimizing the following criteria for global SPRSPR:

Fx=argminF{100(SPR(F)x%)2}F_{x} = arg\min_{F}\left\{ 100 \cdot \left( \text{SPR}(F) - \text{x%} \right)^{2} \right\}

Biological SPRSPR-based reference points are assumed to be regional (Br,xB_{r,x}) and are derived by multiplying SSBPRrfishedSS{BPR}_{r}^{\text{fished}} by regional mean recruitment over a user-defined period:

Br,x=SSBPRrfishedRecYearnyRecAgeRecr,y,snyRecAgeRecYearB_{r,x} = SS{BPR}_{r}^{\text{fished}} \cdot \frac{\sum_{RecYear}^{n_{y} - RecAge}{Rec}_{r,y,s}}{n_{y} - RecAge - RecYear}

Thus, the global SPRSPR calculations result in a global FxF_{x} and regional Br,xB_{r,x} estimates.

Maximum Sustainable Yield (Beverton-Holt)

Similar to deriving spatial reference points for SPRSPR rates, MSYMSY-based reference points assuming a Beverton-Holt stock recruitment relationship can be derived either assuming independent populations without movement or a global population with movement processes incorporated. In cases where density-dependence is defined locally (i.e., area-specific stock-recruitment curves), local MSYMSY-based reference points can also be derived (Kapur et al., 2021).

Independent

In the case where independent spatial regions are assumed, region-specific Fr,msyF_{r,msy} and Br,msyB_{r,msy} estimates can be obtained. In general, all calculations are derived in the same manner as equations described for computing MSYMSY in a single region case, with the exception that demographic rates and fishery selectivity additionally include a region subscript. Notably, in these calculations, virgin recruitment is considered to be regional to compute equilibrium recruitment and is derived in the following manner:

μrRec=μRecζr\mu_{r}^{Rec} = \mu^{Rec}\zeta_{r}

EqRecr=4hrμrRecSSBPRrfishedSSBPRrunfished(1hr)+SSBPRrfished(5hr1){EqRec}_{r} = \frac{4 \cdot h_{r} \cdot \mu_{r}^{Rec} \cdot SS{BPR}_{r}^{\text{fished}}}{SS{BPR}_{r}^{\text{unfished}} \cdot \left( 1 - h_{r} \right) + SS{BPR}_{r}^{\text{fished}} \cdot \left( 5 \cdot h_{r} - 1 \right)}

Fr,msyF_{r,msy} can then be solved for by minimizing (maximizing) yield for each region independently:

Fr,msy=argminFrYieldrF_{r,msy} = arg\min_{F_{r}}\ Yield_{r}

and Br,msyB_{r,msy} can then be written as:

Br,msy=SSBPRrfishedEqRecrB_{r,msy} = SS{BPR}_{r}^{fished}{EqRec}_{r}

Global

In a spatial context, global MSYMSY explicitly accounts for movement dynamics and results in a global FmsyF_{msy} estimate, but regional estiamtes of Br,msyB_{r,msy} because recruitment parameters are defined regionally. Thus, global MSYMSY results in a reference point that achieves rBr,msy\sum_{r}^{}B_{r,msy} for the entire population if FmsyF_{msy} is applied to each region, but does not ensure that each region equilibrates at Br,msyB_{r,msy}.

The following inputs are required for computing global MSYMSY:

  • the relative fishing mortality between fishery fleets (Fratr,f=Fmortr,y=ny,f/fFmortr,y=ny,fFrat_{r,f} = Fmort_{r,y = n_{y},f}/\sum_{f}^{}{Fmort_{r,y = n_{y},f}}),
  • fleet-specific fishery selectivity (Selr,y=y*a,s,fFsh\text{Sel}_{r,y = y^{*}a,s,f}^{\text{Fsh}}),
  • natural mortality for females (Natmortr,y=y*,a,s=1\text{Natmort}_{r,y = y^{*},a,s = 1}),
  • spawning weight at age for females (Wr,y=y*,a,s=1spawnW_{r,y = y^{*},a,s = 1}^{spawn}),
  • maturity at age for females (Matr,y=y*,a,s=1Mat_{r,y = y^{*},a,s = 1}),
  • a matrix of movement values (𝐌y=y*,a,s=1\mathbf{M}_{y = y^{*},\, a,\, s = 1}),
  • spawn timing (tspwnt^{spwn}),
  • the female recruitment sex ratio (ψr,y=y*,s=1\psi_{r,y = y^{*},s = 1}),
  • an estimate of virgin recruitment (μRec\mu^{Rec}),
  • the global steepness value to be used, which is taken as the mean across all regions if steepness is estimated to be regional, consistent with the definitions of global density dependence (h=rhrnrh = \frac{\sum_{r}^{}h_{r}}{n_{r}}), and
  • the recruitment proportions (apportionment) by area (ζr\zeta_{r}).

Using the standard per-recruit computations, global MSYMSY reference points are calculated by first setting the regional numbers-at-age per-recruit equal to the estimated recruitment apportionment parameters (ζr\zeta_{r}) multiplied by the female recruitment sex ratio, which sum to 1:

Nr,a=1fished=ζrψr,y=y*,s=1N_{r,a = 1}^{\text{fished}} = \zeta_{r} \cdot \psi_{r,y = y^{*},s = 1}

Nr,a=1unfished=ζrψr,y=y*,s=1N_{r,a = 1}^{\text{unfished}} = \zeta_{r} \cdot \psi_{r,y = y^{*},s = 1}

After the numbers-at-age per-recruit quantities are initialized, Markovian movement dynamics are applied:

𝐍afished=(𝐍afished)𝐌y=y*,a,s=1for a=amin,,A,amin={1if recruits move2otherwise\mathbf{N}_{a}^{\text{fished}} = \left( \mathbf{N}_{a}^{\text{fished}} \right)^{\top}\mathbf{M}_{y = y^{*},a,s = 1}\quad\text{for }a = a_{\min},\ldots,A,\quad a_{\min} = \left\{ \begin{matrix} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \\ \end{matrix} \right.\

𝐍aunfished=(𝐍aunfished)𝐌y=y*,a,s=1for a=amin,,A,amin={1if recruits move2otherwise\mathbf{N}_{a}^{\text{unfished}} = \left( \mathbf{N}_{a}^{\text{unfished}} \right)^{\top}\mathbf{M}_{y = y^{*},a,s = 1}\quad\text{for }a = a_{\min},\ldots,A,\quad a_{\min} = \left\{ \begin{matrix} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \\ \end{matrix} \right.\

Nr,afishedN_{r,a}^{fished} can then be projected forward as:

Nr,afished=Nr,a1fishedexp[(NatMortr,y=y*,a1,s=1+fFratr,fFmsySelr,y=y*,a1,s=1,fFsh)]if 2<a<AN_{r,a}^{\text{fished}} = {N_{r,a - 1}^{fished} \cdot exp}\begin{matrix} \left\lbrack - \left( NatMort_{r,y = y^{*},a - 1,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{msy} \cdot \text{Sel}_{r,y = y^{*},a - 1,s = 1,f}^{\text{Fsh}} \right) \right\rbrack \\ \text{if }2 < a < A \\ \end{matrix}

For Nr,aunfishedN_{r,a}^{\text{unfished}}, this is given by:

Nr,aunfished=Nr,a1unfishedexp[(NatMortr,y=y*,a1,s=1)],if 2<a<AN_{r,a}^{\text{un}\text{fished}} = {N_{r,a - 1}^{unfished} \cdot exp}\begin{matrix} \left\lbrack - \left( NatMort_{r,y = y^{*},a - 1,s = 1} \right) \right\rbrack,\ \ \ \text{if }2 < a < A \\ \end{matrix}

Baranov’s catch equation is then invoked to compute regional YPRYPR:

Cr,a=fFratr,fFmsySelr,y=y*,a,s=1,fFsh(NatMortr,y=y*,a,s=1+fFratr,fFmsySelr,y=y*,a,s=1,fFsh)Nr,afished(1exp[(NatMortr,y=y*,a,s=1+fFratr,fFmsySelr,y=y*,a,s=1,fFsh)])C_{r,a} = \frac{\sum_{f}^{}\text{Frat}_{r,f} \cdot F_{\text{msy}} \cdot \text{Sel}_{r,y = y^{*},a,s = 1,f}^{\text{Fsh}}}{\left( NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{\text{msy}} \cdot \text{Sel}_{r,y = y^{*},a,s = 1,f}^{\text{Fsh}} \right)} \cdot N_{r,a}^{\text{fished}} \cdot \left( 1 - exp\left\lbrack - \left( NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{\text{msy}} \cdot \text{Sel}_{r,y = y^{*},a,s = 1,f}^{\text{Fsh}} \right) \right\rbrack \right)

Regional equilibrium recruitment is then computed using the Beverton-Holt stock recruitment relationship. To first derive these quantities, regional fished and unfished numbers-at-age per-recruit need to be converted to SSBPR quantities. For regional fished SSBPR (SSBPRrfished)SS{BPR}_{r}^{\text{fished}}), this is written as:

SSBPRrfished=aNr,afishedWr,y=y*,a,s=1spawnMatr,y=y*,a,s=1exp(tspawnNatMortr,y=y*,a,s=1+fFratr,fFmsySelr,y=y*a,s=1,fFsh)SS{BPR}_{r}^{\text{fished}} = \sum_{a}^{}N_{r,a}^{\text{fished}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{r,y = y^{*},a,s = 1} \cdot exp\left( - t^{spawn} \cdot NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{msy} \cdot \text{Sel}_{r,y = y^{*}a,s = 1,f}^{\text{Fsh}} \right)

Regional unfished SSBPRSSBPR (SSBPRrunfished)SS{BPR}_{r}^{\text{unfished}}) is computed in a similar manner:

SSBPRrunfished=aNr,aunfishedWr,y=y*,a,s=1spawnMatr,y=y*,a,s=1exp(tspwnNatMortr,y=y*,a,s=1)SS{BPR}_{r}^{\text{unfished}} = \sum_{a}^{}N_{r,a}^{\text{unfished}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{r,y = y^{*},a,s = 1} \cdot exp\left( - t_{\text{spwn}} \cdot NatMort_{r,y = y^{*},a,s = 1} \right)

Calculations for equilibrium recruitment then follow as:

EqRecr=4hμRecζrrSSBPRrfishedrSSBPRrunfished(1h)+rSSBPRrfished(5h1){EqRec}_{r} = \frac{4 \cdot h \cdot \mu^{Rec}\zeta_{r} \cdot \sum_{r}^{}{SS{BPR}_{r}^{\text{fished}}}}{\sum_{r}^{}{SS{BPR}_{r}^{\text{un}\text{fished}}} \cdot (1 - h) + \sum_{r}^{}{SS{BPR}_{r}^{\text{fished}}} \cdot (5 \cdot h - 1)}

Yield and Br,msyB_{r,msy} are calculated as:

Yield=arCr,aWr,y=y*,a,s=1spawnEqRecrBr,msy=SSBPRrfishedEqRecr{Yield = \sum_{a}^{}{\sum_{r}^{}C_{r,a}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot {EqRec}_{r} }{B_{r,msy} = SS{BPR}_{r}^{\text{fished}} \cdot {EqRec}_{r}}

FmsyF_{msy} can then be solved for using the following:

Fmsy=argminFYieldF_{msy} = arg\min_{F}\ Yield

where the system wide yield is minimized (or maximized).

Local

A key challenge in estimating local spatial reference points (i.e., meta-population dynamics) is that combinations of local fishing mortality rates can sometimes be non-identifiable, because multiple combinations of local reference points can produce similar solutions. However, as highlighted by Kapur et al., 2021, local spatial reference points under density-dependence can potentially be estimated by tracking area-specific yields resulting from a single recruit in each spawning area. This allows the yield surface to be defined and can be used to compute local reference points such as Fr,msyF_{r,msy} and Br,msyB_{r,msy}.

The following inputs are required for computing local MSYMSY:

  • the relative fishing mortality between fishery fleets (Fratr,f=Fmortr,y=ny,f/fFmortr,y=ny,fFrat_{r,f} = Fmort_{r,y = n_{y},f}/\sum_{f}^{}{Fmort_{r,y = n_{y},f}}),
  • fleet-specific fishery selectivity (Selr,y=y*a,s,fFsh\text{Sel}_{r,y = y^{*}a,s,f}^{\text{Fsh}}),
  • natural mortality for females (Natmortr,y=y*,a,s=1\text{Natmort}_{r,y = y^{*},a,s = 1}),
  • spawning weight at age for females (Wr,y=y*,a,s=1spawnW_{r,y = y^{*},a,s = 1}^{spawn}),
  • maturity at age for females (Matr,y=y*,a,s=1Mat_{r,y = y^{*},a,s = 1}),
  • a matrix of movement values (𝐌y=y*,a,s=1\mathbf{M}_{y = y^{*},\, a,\, s = 1}),
  • spawn timing (tspwnt^{spwn}),
  • the female recruitment sex ratio (ψr,y=y*,s=1\psi_{r,y = y^{*},s = 1}),
  • an estimate of global virgin recruitment (μRec\mu^{Rec}),
  • the local steepness value to be used (hrh_{r}),
  • the recruitment proportions (apportionment) by area (ζr\zeta_{r}), which partitions out the estimate of global virgin recruitment.

To ensure identifiability, quantities are tracked by origin region (oo) and destination region (rr). Using standard per-recruit calculations, each region is initialized with one female recruit (i.e., the recruitment sex ratio):

No,r,a=1fished={ψr,y=y*,s=1,ifo=r0,ifor,No,r,a=1unfished={ψr,y=y*,s=1,ifo=r0,iforN_{o,r,a = 1}^{\text{fished}} = \left\{ \begin{matrix} \psi_{r,y = y^{*},s = 1},\ \ \ if o = r \\ 0,\ \ \ if o\ \neq r \\ \end{matrix} \right.\ ,\ \ N_{o,r,a = 1}^{\text{un}\text{fished}} = \left\{ \begin{matrix} \psi_{r,y = y^{*},s = 1},\ \ \ if o = r \\ 0,\ \ \ if o\ \neq r \\ \end{matrix} \right.\

Markovian movement dynamics are then applied:

𝐍o,afished=(𝐍o,afished)𝐌y=y*,a,s=1for a=amin,,A,amin={1if recruits move2otherwise\mathbf{N}_{o,a}^{\text{fished}} = \left( \mathbf{N}_{o,a}^{\text{fished}} \right)^{\top}\mathbf{M}_{y = y^{*},a,s = 1}\quad\text{for }a = a_{\min},\ldots,A,\quad a_{\min} = \left\{ \begin{matrix} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \\ \end{matrix} \right.\

𝐍o,aunfished=(𝐍o,aunfished)𝐌y=y*,a,s=1for a=amin,,A,amin={1if recruits move2otherwise\mathbf{N}_{o,a}^{\text{unfished}} = \left( \mathbf{N}_{o,a}^{\text{unfished}} \right)^{\top}\mathbf{M}_{y = y^{*},a,s = 1}\quad\text{for }a = a_{\min},\ldots,A,\quad a_{\min} = \left\{ \begin{matrix} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \\ \end{matrix} \right.\

No,d,afishedN_{o,d,a}^{fished} can then be projected forward as:

No,r,afished=No,r,a1fishedexp[(NatMortr,y=y*,a1,s=1+fFratr,fFr,msySelr,y=y*,a1,s=1,fFsh)]if 2<a<AN_{o,r,a}^{\text{fished}} = {N_{o,r,a - 1}^{fished} \cdot exp}\begin{matrix} \left\lbrack - \left( NatMort_{r,y = y^{*},a - 1,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{r,msy} \cdot \text{Sel}_{r,y = y^{*},a - 1,s = 1,f}^{\text{Fsh}} \right) \right\rbrack \\ \text{if }2 < a < A \\ \end{matrix}

For No,r,aunfishedN_{o,r,a}^{\text{unfished}}, this is given by:

No,r,aunfished=No,r,a1unfishedexp[(NatMortr,y=y*,a1,s=1)],if 2<a<AN_{o,r,a}^{\text{un}\text{fished}} = {N_{o,r,a - 1}^{unfished} \cdot exp}\begin{matrix} \left\lbrack - \left( NatMort_{r,y = y^{*},a - 1,s = 1} \right) \right\rbrack,\ \ \ \text{if }2 < a < A \\ \end{matrix}

Using Baranov’s catch equation, area-specific YPRYPR can be calculated as:

Co,r,a=fFratr,fFr,msySelr,y=y*,a,s=1,fFsh(NatMortr,y=y*,a,s=1+fFratr,fFr,msySelr,y=y*,a,s=1,fFsh)No,r,afished(1exp[(NatMortr,y=y*,a,s=1+fFratr,fFmsySelr,y=y*,a,s=1,fFsh)])C_{o,r,a} = \frac{\sum_{f}^{}\text{Frat}_{r,f} \cdot F_{r,msy} \cdot \text{Sel}_{r,y = y^{*},a,s = 1,f}^{\text{Fsh}}}{\left( NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{r,msy} \cdot \text{Sel}_{r,y = y^{*},a,s = 1,f}^{\text{Fsh}} \right)} \cdot N_{o,r,a}^{\text{fished}} \cdot \left( 1 - exp\left\lbrack - \left( NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{\text{msy}} \cdot \text{Sel}_{r,y = y^{*},a,s = 1,f}^{\text{Fsh}} \right) \right\rbrack \right)

Regional equilibrium recruitment in each destination is computed by solving a non-linear Beverton-Holt stock-recruitment relationship that ensures internal consistency. To derive these quantities, regional fished and unfished numbers-at-age per-recruit are converted to SSBPR quantities. For regional fished SSBPR (SSBPRo,rfished)SS{BPR}_{o,r}^{\text{fished}}), this is written as:

SSBPRo,rfished=aNo,r,afishedWr,y=y*,a,s=1spawnMatr,y=y*,a,s=1exp(tspawnNatMortr,y=y*,a,s=1+fFratr,fFmsySelr,y=y*a,s=1,fFsh)SS{BPR}_{o,r}^{\text{fished}} = \sum_{a}^{}N_{o,r,a}^{\text{fished}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{r,y = y^{*},a,s = 1} \cdot exp\left( - t^{spawn} \cdot NatMort_{r,y = y^{*},a,s = 1} + \sum_{f}^{}\text{Frat}_{r,f} \cdot F_{msy} \cdot \text{Sel}_{r,y = y^{*}a,s = 1,f}^{\text{Fsh}} \right)

Regional unfished SSBPRSSBPR (SSBPRo,runfished)SS{BPR}_{o,r}^{\text{unfished}}) is computed in a similar manner:

SSBPRo,runfished=aNo,r,aunfishedWr,y=y*,a,s=1spawnMatr,y=y*,a,s=1exp(tspwnNatMortr,y=y*,a,s=1)SS{BPR}_{o,r}^{\text{unfished}} = \sum_{a}^{}N_{o,r,a}^{\text{unfished}} \cdot W_{r,y = y^{*},a,s = 1}^{spawn} \cdot \text{Mat}_{r,y = y^{*},a,s = 1} \cdot exp\left( - t_{\text{spwn}} \cdot NatMort_{r,y = y^{*},a,s = 1} \right)

Next, define the components of a Beverton-Holt stock recruitment function:

Ar=4hrζrμRecA_{r} = 4 \cdot h_{r} \cdot \zeta_{r} \cdot \mu^{Rec}

Br=(1hr)oSSBPRo,runfishedζrμRecB_{r} = \left( 1 - h_{r} \right) \cdot \ \sum_{o}^{}{SS{BPR}_{o,r}^{\text{unfished}} \cdot \zeta_{r} \cdot \mu^{Rec}}

Dr=5hr1D_{r} = 5 \cdot h_{r} - 1

Equilibrium recruitment in the destination region (DestEqRecr{DestEqRec}_{r}) is then defined as a function of contributions from all origin regions (OrigEqReco{OrigEqRec}_{o}):

DestEqRecr=g(OrigEqReco)=ArEqSSBrfishedBr+DrEqSSBrfishedEqSSBrfished=oSSBPRo,rfishedOrigEqReco{{DestEqRec}_{r} = g({OrigEqRec}_{o}) = \frac{A_{r} \cdot EqSSB_{r}^{\text{fished}}}{B_{r} + D_{r} \cdot EqSSB_{r}^{\text{fished}}} }{EqSSB_{r}^{\text{fished}} = \sum_{o}^{}{SS{BPR}_{o,r}^{\text{fished}} \cdot {OrigEqRec}_{o}}}

where EqSSBrfishedEqSSB_{r}^{\text{fished}} is the fished equilibrium SSBSSB in destination rr generated by contributions from all origin regions. The aforementioned equation is a non-linear system and does not have an analytical solution. Thus, an iterative solution is obtained using Newton-Raphson’s method. Although the function is expressed in terms of destination recruitment, it is the origin recruitment that is iteratively solved for. Specifically, Newton-Raphson’s method adjusts OrigEqReco{OrigEqRec}_{o} until the contributions from all origins result in self-consistent DestEqRecr{DestEqRec}_{r} values.

Total yield (YieldYield) and yield in the destination region (Yieldr{Yield}_{r}) is then defined as:

Yieldr=o(aCo,r,aWr,y=y*,a,s=1spawn)OrigEqRecoYield=rYieldr{{Yield}_{r} = \sum_{o}^{}{\left( \sum_{a}^{}{C_{o,r,a}W_{r,y = y^{*},a,s = 1}^{spawn}} \right){OrigEqRec}_{o}} }{Yield = \sum_{r}^{}{Yield}_{r}}

Br,msy=oSSBPRo,rfishedOrigEqRecoB_{r,msy} = \sum_{o}^{}{SS{BPR}_{o,r}^{\text{fished}} \cdot {OrigEqRec}_{o}}

Lastly, regional estimates of Fr,msyF_{r,msy} are solved for by minimizing total yield:

Fr,msy=argminFr,msyYieldF_{r,msy} = \arg\min_{F_{r,msy}}\ Yield

where the system wide yield is minimized (or maximized).

Deriving Catch Advice and Projections

A core part of the assessment process is to convert reference point estimates into catch advice. In the following sections, we will mathematically describe how catch advice is derived, and proceed to provide code examples for demonstration. To conduct projection from the terminal year, users must define the following quantities:

  • Terminal year estimates of numbers-at-age,
  • A user defined period of of recruitment values to use,
  • A user defined period of weight-at-age values to use for projections,
  • A user defined period of natural mortality-atage values to use for projections,
  • A user defined period of maturity-at-age values to use for projections,
  • A user defined period of fishery selectivity values to use for projections,
  • A user defined period of movement values to use for projections,
  • Terminal year estimates of fishing mortality,
  • Fishing mortality rate to use to decrement the population,

Optionally, users can define:

  • Biological reference points to use to project fishing mortality in the subsequent years, if a harvest control rule is utilized, and
  • A function describing a harvest control rule.

In the first year of the projection period, projected fishing mortality is determined with estimates of fishing mortality in the terminal year of the assessment:

projFr,y=fFmortr,y=term,f projF_{r,y} = \sum_f Fmort_{r,y = term,f}

Total mortality can then be computed as:

Fr,y,a,s,f=projFr,yFratr,fSelr,y,a,s,fFsh F_{r,y,a,s,f} = projF_{r,y} \cdot Frat_{r,f} \cdot Sel^{Fsh}_{r,y,a,s,f}

Zr,y,a,s,f=fFr,y,a,s,f+NatMortr,y,a,s Z_{r,y,a,s,f} = \sum_f F_{r,y,a,s,f} + NatMort_{r,y,a,s}

Similarly, projected numbers at age in the first year utilizes estimates of numbers at age in the terminal year of the assessment, for which movement has already been applied. An exponential mortality model is then used to determine the numbers at age in the next year (y+1y+1):

projNr,y,a,s=Nr,y=term,a,s projN_{r,y,a,s} = N_{r,y = term,a,s}

projNr,y+1,a+1,s=projNr,y,a,sexp(Zr,y,a,s),for 1a<a+projNr,y+1,a,s=projNr,y,a1,sexp(Zr,y,a1,s)+projNr,y,a,sexp(Zr,y,a,s),for a=a+ \begin{aligned} projN_{r,y+1,a+1,s} = projN_{r,y,a,s}\exp(-Z_{r,y,a,s}), \quad \text{for } 1 \leq a < a_+ \\ projN_{r,y+1,a,s} = projN_{r,y,a-1,s}\exp(-Z_{r,y,a-1,s}) + projN_{r,y,a,s}\exp(-Z_{r,y,a,s}), \quad \text{for } a = a_+ \end{aligned}

Quantities of spawning stock biomass can then be computed as:

projSSBr,y=aa+projNr,y,a,s=1Wr,y,a,s=1Matr,y,a,s=1 projSSB_{r,y} = \sum_a^{a+} projN_{r,y,a,s=1}W_{r,y,a,s=1}Mat_{r,y,a,s=1}

Additionally, quantities of projected catch can be derived using Baranov’s catch equation:

projCr,y,a,s,fa=Fmortr,y,fSelr,y,a,s,fFshZr,y,a,sprojNr,y,a,s[1exp(Zr,y,a,s)] projC^a_{r,y,a,s,f} = \frac{Fmort_{r,y,f}Sel^{Fsh}_{r,y,a,s,f}}{Z_{r,y,a,s}} projN_{r,y,a,s} \left[1-\exp(-Z_{r,y,a,s})\right]

projCatchr,y,f=aa+snsprojCr,y,a,s,faWr,y,a,s projCatch_{r,y,f} = \sum_a^{a_+} \sum_s^{n_s} projC^a_{r,y,a,s,f} W_{r,y,a,s}

Fishing mortality in the next year can then be projected forward using either a harvest control rule, or projected forward using user inputs:

projFr,y+1={f(SSBr,y;BRPr,y;FRPr,y)if using a HCRf(Finputr,y)if using a user defined fishing mortality rate projF_{r,y+1} = \begin{cases} f(SSB_{r,y}; BRP_{r,y}; FRP_{r,y}) & \text{if using a HCR} \\ f(Finput_{r,y}) & \text{if using a user defined fishing mortality rate} \end{cases}

where f()f() is a harvest control function that takes the inputs SSBSSB, BRPBRP (biological reference points), and FRPFRP (a fishery reference point). Alternatively, f()f() can be a user defined matrix of fishing mortality rates to use during the projection period across regions. Projected fishing mortality is then summed with natural mortaltiy to compute the projected total mortality in a given projection year.

Recruitment dynamics are then projected in each year following the initial projection year. In particular, several recruitment projection options are availiable. These include both deterministic predictions as well as the ability to incorporate stochasticity into recruitment projections.

In particular, deterministic recruitment has the option to be projected forward as zero:

projNr,y,1,s=0,if y > 1 projN_{r,y,1,s} = 0, \text{if y > 1}

where no recruitment occurs. Deterministic recruitment can also be projected forward using mean recruitment (mean_rec) from a matrix of estimated recruitment values from the assessment model supplied by the user:

projNr,y,1,s=yRecr,yn,if y > 1 projN_{r,y,1,s} = \frac{\sum_y Rec_{r,y}}{n}, \text{if y > 1}

Alternatively, users can also specify a Beverton-Holt stock recruitment function (bh_rec) to be used for deterministic recruitment projection, which then requires users to supply the necessary parameter inputs. In the case where local recruitment is specified, this is computed as (i.e., metapopulation dynamics):

projNr,y,1,s=4hrRr,0projSSBrSSBrunfished(1hr)+projSSBr(5hr1) projN_{r,y,1,s} = \frac{4\cdot h_r \cdot R_{r,0} \cdot projSSB_r}{SSB_r^{\text{unfished}} \cdot (1 - h_r) + projSSB_r \cdot (5 \cdot h_r - 1)}

By contrast, if global recruitment is specified, this is computed as:

projNr,y,1,s=(4hR0rprojSSBrrSSBrunfished(1h)+rprojSSBr(5h1))ζr projN_{r,y,1,s} = \left( \frac{4\cdot h \cdot R_{0} \cdot \sum_r projSSB_r}{\sum_r SSB_r^{\text{unfished}} \cdot (1 - h) + \sum_r projSSB_r \cdot (5 \cdot h - 1)} \right) \zeta_r

where density-dependence occurs globally, and a recruitment apportionment parameter is utilized to partition global recruits in a given year.

Lastly, users can specify recruitment projections to be stochastic, wherein an inverse Gaussian (inv_gauss) distribution parameterized based on estimated recruitment values from the assessment model is utilized to project recruitment into the future:

AMeanRecr=1Yy=avgYRecr,y \text{AMeanRec}_r = \frac{1}{Y} \sum_{y = avg}^{Y} \text{Rec}_{r,y}

HMeanRecr=(1Yy=avgY1Recr,y)1 \text{HMeanRec}_r = \left( \frac{1}{Y} \sum_{y = avg}^{Y} \frac{1}{\text{Rec}_{r,y}} \right)^{-1}

γr=AMeanRecrHMeanRecr \gamma_r = \frac{\text{AMeanRec}_r}{\text{HMeanRec}_r}

βr=AMeanRecr \beta_r = \text{AMeanRec}_r

δr=1γr1 \delta_r = \frac{1}{\gamma_r - 1}

CVRecr=1δr \text{CVRec}_r = \sqrt{ \frac{1}{\delta_r} }

For each year a random draw is made from a standard normal distribution, which is then transformed:

ψy=By2,where ByN(0,1) \psi_y = B_y^2, \quad \text{where } B_y \sim N(0,1)

ωr,y=βr(1+ψy4δrψy+ψy22δr) \omega_{r,y} = \beta_r \left( 1 + \frac{\psi_y - \sqrt{4 \delta_r \psi_y + \psi_y^2}}{2 \delta_r} \right)

ζr,y=βr(1+ψy+4δrψy+ψy22δr) \zeta_{r,y} = \beta_r \left( 1 + \frac{\psi_y + \sqrt{4 \delta_r \psi_y + \psi_y^2}}{2 \delta_r} \right)

θr,y=βrβr+ωr,i \theta_{r,y} = \frac{\beta_r}{\beta_r + \omega_{r,i}}

Then, a draw is conducted U(0,1)\sim U(0,1), and simulated recruitment is defined as:

Recr,ysim={ωr,y,if Uyθr,yζr,y,otherwise \text{Rec}_{r,y}^{\text{sim}} = \begin{cases} \omega_{r,y}, & \text{if } U_y \leq \theta_{r,y} \\ \zeta_{r,y}, & \text{otherwise} \end{cases}

Thus, this inverse gaussian mixture ensures the simulated values have approximately the correct mean and variability based on historical recruitment values.

After recruitment processes occur, the usual dynamics are then applied to project the population forward. Here, movement of individuals first occurs:

𝐩𝐫𝐨𝐣𝐍a=(𝐩𝐫𝐨𝐣𝐍a)𝐌y,a,sfor a=amin,,A,amin={1if recruits move2otherwise \boldsymbol{projN}_a^ = (\boldsymbol{projN}_a)^\top \boldsymbol{M}_{y, a, s} \quad \text{for } a = a_{\min}, \dots, A, \quad a_{\min} = \begin{cases} 1 & \text{if recruits move} \\ 2 & \text{otherwise} \end{cases}

Following movement, which is only applied in projected years y>1y > 1, as it has already been accounted for in the terminal year estimates of numbers at age, the exponential mortality model is applied. Projected spawning stock biomass and catch are then derived. Fishing mortality in subsequent years is updated accordingly. This sequence of steps is repeated until the specified number of projection years is reached. Thus, catch advice for the year following the terminal assessment year corresponds to the projected catch in projection year 2 (i.e., projCatchr,y=2,fprojCatch_{r, y = 2, f}).

Code Demonstration

In the subsequent sections, we will demonstrate how reference points, catch projections, catch advice, and stochastic projections can be derived and conducted using SPoRC. These features rely on users to have a report file from a SPoRC model, and we have generally coded this in a way that there is flexibility for users to define how projections are done.

Getting Reference Points

Single Region

To illustrate how reference points are derived, we begin by extracting the report file from the single-region sablefish case study (sgl_rg_sable_rep). We then call the Get_Reference_Points function to calculate the reference points. In the example below, we estimate the F40F_{40} and B40B_{40} values in a single-region context. This requires passing the sablefish data file (sgl_reg_sable_data) to the data argument, the report file to the rep argument, and setting the SPR rate (SPR_x) to 0.4. We also specify that the reference point is SPR-based and pertains to a single region. Additional inputs include the first year of recruitment used for calculating B40B_{40}, the recruitment age (which excludes the last rec_age years when computing the mean), the timing of spawning, and the sex ratio used in the B40B_{40} calculation. Note that the sablefish example does not utilize a stock recruitment relationship. However, if a Beverton-Holt stock recruitment relationship is utilized and users want to estimate MSY-based reference points, this can be derived by setting what = 'BH_MSY'.

data("sgl_rg_sable_rep") # read in single region report
data("sgl_rg_sable_data") # read in single region data 

# single area model
sgl_ref_pt <- SPoRC::Get_Reference_Points(data = sgl_rg_sable_data, # data file
                                          rep = sgl_rg_sable_rep, # report file
                                          SPR_x = 0.4, # spr rate
                                          type = 'single_region', # single region reference point
                                          what = 'SPR', # SPR reference point
                                          calc_rec_st_yr = 20, # first year to calculate mean recruitment
                                          rec_age = 2,  # exclues the last rec_age years when computing mean recruitment
                                          t_spwn = 0,  # spawn timing
                                          sex_ratio_f = 0.5 # recruitment sex-ratio for females
                                          )
sgl_ref_pt$f_ref_pt # F40
#> [1] 0.08625413
sgl_ref_pt$b_ref_pt # B40
#> [1] 121.3814

Note that the sablefish example does not utilize a stock recruitment relationship. However, if a Beverton-Holt stock recruitment relationship is utilized and users want to estimate MSY-based reference points, this can be derived by setting what = 'BH_MSY'.

Multi Region

In the following, we will demonstrate how spatial reference points can be derived. In general, this is similar to the single region case, except that a spatial model and associated report files will be needed (report file needs to be spatially-explicit). Again, we will use the five-region sablefish case study as an example, where we will estimate both independent SPR rates and global SPR rates. In contrast to the single region case, type would now need to be specified as multi_region. Additionally, for independent SPR rates where movement dynamics are ignored, what is now set at independent_SPR. All the other arguments are defined the same as the example above. Given that these are treated as independent populations, fishery reference points and biological reference points are region-specific and can be applied accordingly.

data("mlt_rg_sable_rep") # read in multi region report
data("mlt_rg_sable_data") # read in multi region data 

# multi region model with independent SPR
mlt_ref_pt_indp <- Get_Reference_Points(data = mlt_rg_sable_data, # data file
                                          rep = mlt_rg_sable_rep, # report file
                                          SPR_x = 0.4, # spr rate
                                          type = 'multi_region', # multi region reference point
                                          what = 'local_BH_MSY', # SPR reference point
                                          calc_rec_st_yr = 20, # first year to calculate mean recruitment
                                          rec_age = 2,  # exclues the last rec_age years when computing mean recruitment
                                          t_spwn = 0,  # spawn timing
                                          sex_ratio_f = rep(0.5, mlt_rg_sable_data$n_regions) # recruitment sex-ratio for females
                                          )
mlt_ref_pt_indp$f_ref_pt # F40
#> [1] 1.868139e-11 2.913135e-11 1.757183e-10 4.829113e-02 5.021245e-01
mlt_ref_pt_indp$b_ref_pt # B40
#> [1]  7.067837 11.328148  7.138283 32.894687 15.427392

By contrast, users can also specify global SPR rates. This involves simply changing the what argument to global_SPR, which results in a single F40F_{40} being estimated, but region-specific Br,40B_{r,40} given that regional estimates of recruitment are utilized. Note that the F40F_{40} outputs 5 values for the 5 regions modelled in the case study, but these values are all identical.

data("mlt_rg_sable_rep") # read in multi region report
data("mlt_rg_sable_data") # read in multi region data 

# multi region model with global SPR
mlt_ref_pt_global <- SPoRC::Get_Reference_Points(data = mlt_rg_sable_data, # data file
                                                 rep = mlt_rg_sable_rep, # report file
                                                 SPR_x = 0.4, # spr rate
                                                 type = 'multi_region', # multi region reference point
                                                 what = 'global_SPR', # SPR reference point
                                                 calc_rec_st_yr = 20, # first year to calculate mean recruitment
                                                 rec_age = 2,  # exclues the last rec_age years when computing mean recruitment
                                                 t_spwn = 0,  # spawn timing
                                                 sex_ratio_f = rep(0.5, mlt_rg_sable_data$n_regions) # recruitment sex-ratio for females
                                                 )
mlt_ref_pt_global$f_ref_pt # F40
#> [1] 0.08099479 0.08099479 0.08099479 0.08099479 0.08099479
mlt_ref_pt_global$b_ref_pt # B40
#> [1] 31.93313 26.89321 20.22807 30.01715 14.03777

Similarly, MSY-based reference points assuming a Beverton-Holt stock recruitment function can be specified as well. This can be easily specified and involves either assuming independent populations (what = 'independent_BH_MSY') or a population with global density dependence (what = 'global_BH_MSY').

Conducting Catch Projections to Derive Catch Advice (Deterministic Recruitment)

Single Region

Next, using the reference points derived from the single region case study, we can conduct population and catch projections to derive catch advice. Note that this will require users to have a data file to extract the relevant demographic rates and data components, as well as a report file to extract necessary estimates to conduct projections. Let us first define a threshold harvest control rule to utilize in our population projections, although note that this is not strictly necessary.

# Define HCR to use
HCR_function <- function(x, frp, brp, alpha = 0.05) {
  stock_status <- x / brp # define stock status
  # If stock status is > 1
  if(stock_status >= 1) f <- frp
  # If stock status is between brp and alpha
  if(stock_status > alpha && stock_status < 1) f <- frp * (stock_status - alpha) / (1 - alpha)
  # If stock status is less than alpha
  if(stock_status < alpha) f <- 0
  return(f)
}

# Create a tibble for plotting
hcr_df <- tibble(
  i = 1:200,
  SSB_B40 = i / sgl_ref_pt$b_ref_pt,
  F = sapply(i, function(x) {
    HCR_function(x = x, frp = sgl_ref_pt$f_ref_pt, brp = sgl_ref_pt$b_ref_pt)
  })
)

# Plot
ggplot(hcr_df, aes(x = SSB_B40, y = F)) +
  geom_line(color = "steelblue", size = 1) +
  labs(x = "SSB / B40", y = "F") +
  theme_bw(base_size = 13)

We can define all the inputs needed to run the population projection:


data("sgl_rg_sable_rep") # read in single region report
data("sgl_rg_sable_data") # read in single region data 

# Setup necessary inputs
t_spawn <- 0 # spawn timing
n_proj_yrs <- 15 # number of projection years
n_regions <- 1 # number of regions
n_ages <- length(sgl_rg_sable_data$ages) # number of ages
n_sexes <- sgl_rg_sable_data$n_sexes # number of sexes
n_fish_fleets <- sgl_rg_sable_data$n_fish_fleets # number of fishery fleets
do_recruits_move <- 0 # recruits don't move
terminal_NAA <- array(sgl_rg_sable_rep$NAA[,length(sgl_rg_sable_data$years),,], dim = c(n_regions, n_ages, n_sexes)) # terminal numbers at age
terminal_NAA0 <- array(sgl_rg_sable_rep$NAA0[,length(sgl_rg_sable_data$years),,], dim = c(n_regions, n_ages, n_sexes)) # terminal numbers at age
WAA <- array(rep(sgl_rg_sable_data$WAA[,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # weight at age
WAA_fish <- array(rep(sgl_rg_sable_data$WAA[,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes, n_fish_fleets)) # weight at age for fishery
MatAA <- array(rep(sgl_rg_sable_data$MatAA[,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # maturity at age
fish_sel <- array(rep(sgl_rg_sable_rep$fish_sel[,length(sgl_rg_sable_data$years),,,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes, n_fish_fleets)) # selectivity
Movement <- array(rep(sgl_rg_sable_rep$Movement[,,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_regions, n_proj_yrs, n_ages, n_sexes)) # movement
terminal_F <- array(sgl_rg_sable_rep$Fmort[,length(sgl_rg_sable_data$years),], dim = c(n_regions, n_fish_fleets)) # terminal F
natmort <- array(sgl_rg_sable_rep$natmort[,length(sgl_rg_sable_data$years),,], dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # natural mortality
recruitment <- array(sgl_rg_sable_rep$Rec[,20:(length(sgl_rg_sable_data$years) - 2)], dim = c(n_regions, length(20:length(sgl_rg_sable_data$years) - 2))) # recruitment values to use for mean recruitment calculations or inverse gaussian parameterization
sexratio <- array(0.5, dim = c(n_regions, n_proj_yrs, n_sexes)) # recruitment sex ratio 

# Define reference points to use in HCR
f_ref_pt = array(sgl_ref_pt$f_ref_pt, dim = c(n_regions, n_proj_yrs))
b_ref_pt = array(sgl_ref_pt$b_ref_pt, dim = c(n_regions, n_proj_yrs))

Note that in these projections, all demographic rates (e.g., weight-at-age, movement, maturity) use estimates from the terminal year of the assessment. However, this is not required - users may instead define demographic rates for the projection period using other approaches (e.g., averages over the last 5 years). Regardless of the approach, rates must be provided by the user in the format and dimensions described above. A population projection can then be conducted with the Do_Population_Projection function:

# do population projection
out <- Do_Population_Projection(n_proj_yrs = n_proj_yrs, # Number of projection years
                               n_regions = n_regions, # number of regions
                               n_ages = n_ages, # number of ages
                               n_sexes = n_sexes, # number of sexes
                               sexratio = sexratio, # sex ratio for recruitment
                               n_fish_fleets = n_fish_fleets, # number of fishery fleets
                               do_recruits_move = do_recruits_move, # whether recruits move (not used since single area)
                               recruitment = recruitment, # recruitment values to use for mean recruitment 
                               terminal_NAA = terminal_NAA, # terminal numbers at age
                               terminal_NAA0 = terminal_NAA0, 
                               terminal_F = terminal_F, # terminal F
                               natmort = natmort, # natural mortality values to use in projection
                               WAA = WAA, # weight at age values to use in projection spawning
                               WAA_fish = WAA_fish, # weight at age values to use in projection fishery
                               MatAA = MatAA, # maturity at age values to use in projection
                               fish_sel = fish_sel, # fishery selectivity values to use in projection
                               Movement = Movement, # movement values (not used since single area)
                               f_ref_pt = f_ref_pt, # fishery reference points (f40)
                               b_ref_pt = b_ref_pt, # biological reference points (b40)
                               HCR_function = HCR_function, # threshold control rule defined above
                               recruitment_opt = "mean_rec", # recruitment assumption utilizes the mean recruits for the supplied recruitment estimates
                               fmort_opt = "HCR", # Fishing mortality in projection years are determined using a HCR
                               t_spawn = t_spawn # Spawn timing
                               )

The outputted object from the function then includes the projected fishing mortality rates, the projected catch (i.e., the catch advice), the projected spawning stock biomass, the projected numbers at age, and the projected total mortality at age. We can plot a few of these quantities out below. In the example below, we show the projected SSB:

combined_ssb <- c(sgl_rg_sable_rep$SSB[1, -65], out$proj_SSB[1,]) # removing terminal year becauase repeated in projection calculations
years <- 1960:(2023 + n_proj_yrs)

ssb_df <- tibble(
  Year = years,
  SSB = combined_ssb
)

# Plot
ggplot(ssb_df, aes(x = Year, y = SSB)) +
  geom_line(size = 1) +
  geom_vline(xintercept = 2024, linetype = "dashed") + # projection start
  scale_y_continuous(limits = c(0, 300)) +
  labs(x = "Year", y = "SSB (kt)") +
  theme_bw(base_size = 13)

as well as projected catches, which can then be the basis of management advice:

combined_catch <- c(
  rowSums(sgl_rg_sable_rep$PredCatch[1, -65, ]), # removing terminal year becauase repeated in projection calculations
  rowSums(out$proj_Catch[1, , ])
)

years <- 1960:(2023 + n_proj_yrs)

catch_df <- tibble(
  Year = years,
  Catch = combined_catch
)

# Plot
ggplot(catch_df, aes(x = Year, y = Catch)) +
  geom_line(size = 1) +
  geom_vline(xintercept = 2024, linetype = "dashed") +  # projection start
  labs(x = "Year", y = "Catch (kt)") +
  theme_bw(base_size = 13)
sum(out$proj_Catch[1,2,]) # Catch advice in terminal year + 1

Importantly, catch advice should be based on terminal year+1 rather than the first projection year, since the first projection year serves only as an initialization step for the projection.

Multi Region

In the following, we will then demonstrate how catch projections can be conducted in a multi-region context, using independent SPR rates, such that there are region-specific estimates of Fr,40F_{r,40} and Br,40B_{r,40}. In general, the steps are similar to the single-region case. Again, we will utilize the harvest control rule function defined above. Given that each region has their own unique estimates, this will result in different harvest control rules being applied to each region:

# Define HCR to use
HCR_function <- function(x, frp, brp, alpha = 0.05) {
  stock_status <- x / brp # define stock status
  # If stock status is > 1
  if(stock_status >= 1) f <- frp
  # If stock status is between brp and alpha
  if(stock_status > alpha && stock_status < 1) f <- frp * (stock_status - alpha) / (1 - alpha)
  # If stock status is less than alpha
  if(stock_status < alpha) f <- 0
  return(f)
}

# Build a dataframe by looping over j and i scalars
hcr_df <- expand.grid(
  j = 1:5,
  i = 1:50
) %>%
  mutate(
    frp = mapply(function(j) mlt_ref_pt_indp$f_ref_pt[j], j),
    brp = mapply(function(j) mlt_ref_pt_indp$b_ref_pt[j], j),
    F = mapply(function(i, j) {
      HCR_function(x = i, frp = mlt_ref_pt_indp$f_ref_pt[j], brp = mlt_ref_pt_indp$b_ref_pt[j])
    }, i, j),
    SSB_B40 = i / brp
  )

ggplot(hcr_df, aes(x = SSB_B40, y = F, color = factor(j))) +
  geom_line(lwd = 1.3) +
  facet_wrap(~j, scales = 'free') +
  labs(x = "SSB / B40", y = "F",  color = 'Region') +
  theme_bw(base_size = 13) +
  theme(legend.position = 'none')

Let’s then define all the inputs needed to run the population projection:


data("mlt_rg_sable_rep") # read in multi region report
data("mlt_rg_sable_data") # read in multi region data 

# Setup necessary inputs
t_spawn <- 0 # spawn timing
n_proj_yrs <- 15 # number of projection years
n_regions <- 5 # number of regions
n_ages <- length(mlt_rg_sable_data$ages) # number of ages
n_sexes <- mlt_rg_sable_data$n_sexes # number of sexes
n_fish_fleets <- mlt_rg_sable_data$n_fish_fleets # number of fishery fleets
do_recruits_move <- 0 # recruits don't move
terminal_NAA <- array(mlt_rg_sable_rep$NAA[,length(mlt_rg_sable_data$years),,], dim = c(n_regions, n_ages, n_sexes)) # terminal numbers at age
terminal_NAA0 <- array(mlt_rg_sable_rep$NAA0[,length(mlt_rg_sable_data$years),,], dim = c(n_regions, n_ages, n_sexes)) # terminal numbers at age
WAA <- array(rep(mlt_rg_sable_data$WAA[,length(mlt_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # weight at age
WAA_fish <- array(rep(mlt_rg_sable_data$WAA[,length(mlt_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes, n_fish_fleets)) # weight at age
MatAA <- array(rep(mlt_rg_sable_data$MatAA[,length(mlt_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # maturity at age
fish_sel <- array(rep(mlt_rg_sable_rep$fish_sel[,length(mlt_rg_sable_data$years),,,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes, n_fish_fleets)) # selectivity
Movement <- aperm(abind::abind(replicate(n_proj_yrs, mlt_rg_sable_rep$Movement[,,length(mlt_rg_sable_data$years),,], simplify = FALSE), along = 5), perm = c(1,2,5,3,4)) # movement
terminal_F <- array(mlt_rg_sable_rep$Fmort[,length(mlt_rg_sable_data$years),], dim = c(n_regions, n_fish_fleets)) # terminal F
natmort <- array(mlt_rg_sable_rep$natmort[,length(mlt_rg_sable_data$years),,], dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # natural mortality
recruitment <- array(mlt_rg_sable_rep$Rec[,20:(length(mlt_rg_sable_data$years) - 2)], dim = c(n_regions, length(20:length(mlt_rg_sable_data$years) - 2))) # recruitment values to use for mean recruitment calculations or inverse gaussian parameterization
sexratio <- array(0.5, dim = c(n_regions, n_proj_yrs, n_sexes)) # recruitment sex ratio 

# Define independent SPR reference points to use in HCR
f_ref_pt_indp = array(mlt_ref_pt_indp$f_ref_pt, dim = c(n_regions, n_proj_yrs))
b_ref_pt_indp = array(mlt_ref_pt_indp$b_ref_pt, dim = c(n_regions, n_proj_yrs))

When global reference points (SPR or MSY-based) are used, the same fishing mortality (FF) is applied across all regions because only a single global FF is estimated. This is illustrated below:

f_ref_pt_global = array(mlt_ref_pt_global$f_ref_pt, dim = c(n_regions, n_proj_yrs))
b_ref_pt_global = array(mlt_ref_pt_global$b_ref_pt, dim = c(n_regions, n_proj_yrs))

By contrast, the independent SPR approach assigns a region-specific FF value. For example, in the first projection year:

# F40
f_ref_pt_indp[,2] # independent SPR
#> [1] 1.868139e-11 2.913135e-11 1.757183e-10 4.829113e-02 5.021245e-01
f_ref_pt_global[,2] # global SPR
#> [1] 0.08099479 0.08099479 0.08099479 0.08099479 0.08099479

# b40
b_ref_pt_indp[,2] # independent SPR
#> [1]  7.067837 11.328148  7.138283 32.894687 15.427392
b_ref_pt_global[,2] # global SPR
#> [1] 31.93313 26.89321 20.22807 30.01715 14.03777

For the projections that follow, we use independent SPR rates to allow for region-specific reference points:

# do population projection
out <- Do_Population_Projection(n_proj_yrs = n_proj_yrs, # Number of projection years
                              n_regions = n_regions, # number of regions
                              n_ages = n_ages, # number of ages
                              n_sexes = n_sexes, # number of sexes
                              sexratio = sexratio, # sex ratio for recruitment
                              n_fish_fleets = n_fish_fleets, # number of fishery fleets
                              do_recruits_move = do_recruits_move, # whether recruits move (not used since single area)
                              recruitment = recruitment, # recruitment values to use for mean recruitment 
                              terminal_NAA = terminal_NAA, # terminal numbers at age
                              terminal_NAA0 = terminal_NAA0,
                              terminal_F = terminal_F, # terminal F
                              natmort = natmort, # natural mortality values to use in projection
                              WAA = WAA, # weight at age values to use in projection spawning
                              WAA_fish = WAA_fish, # weight at age values to use in projection fishery
                              MatAA = MatAA, # maturity at age values to use in projection
                              fish_sel = fish_sel, # fishery selectivity values to use in projection
                              Movement = Movement, # movement values (not used since single area)
                              f_ref_pt = f_ref_pt_indp, # fishery reference points (f40)
                              b_ref_pt = b_ref_pt_indp, # biological reference points (b40)
                              HCR_function = HCR_function, # threshold control rule defined above
                              recruitment_opt = "mean_rec", # recruitment assumption utilizes the mean recruits for the supplied recruitment estimates
                              fmort_opt = "HCR", # Fishing mortality in projection years are determined using a HCR
                              t_spawn = t_spawn # Spawn timing
                              )

Again, we can visualize what these projections look like in terms of SSB and catch advice:

combined_ssb <- cbind(mlt_rg_sable_rep$SSB[,-62], out$proj_SSB[,]) # removing terminal year becauase repeated in projection calculations
combined_ssb_df <- reshape2::melt(combined_ssb) %>% 
  rename(Region = Var1, Year = Var2, SSB = value)

# Plot
ggplot(combined_ssb_df, aes(x = Year + 1959, y = SSB, color = factor(Region))) +
  geom_line(size = 1) +
  geom_vline(xintercept = 2021, linetype = "dashed") + # projection start
  facet_wrap(~Region) +
  scale_y_continuous(limits = c(0, NA)) +
  labs(x = "Year", y = "SSB (kt)") +
  theme_bw(base_size = 13) +
  theme(legend.position = 'none')

combined_catch <- cbind(apply(mlt_rg_sable_rep$PredCatch, c(1,2), sum), apply(out$proj_Catch, c(1,2), sum))
combined_catch_df <- reshape2::melt(combined_catch) %>% 
  rename(Region = Var1, Year = Var2, Catch = value)

# Plot
ggplot(combined_catch_df, aes(x = Year + 1959, y = Catch, color = factor(Region))) +
  geom_line(size = 1) +
  geom_vline(xintercept = 2021, linetype = "dashed") + # projection start
  facet_wrap(~Region) +
  scale_y_continuous(limits = c(0, NA)) +
  labs(x = "Year", y = "Catch") +
  theme_bw(base_size = 13) +
  theme(legend.position = 'none')

rowSums(out$proj_Catch[,2,]) # Catch advice by region in terminal year + 1

Conducting Stochastic Population Projections

In the final section of this vignette, we demonstrate how to conduct stochastic population projections. For simplicity, we focus on the single-region case, though the approach extends similarly to multi-region scenarios. Stochastic projections follow the same general structure as deterministic ones, with the key difference being that recruitment_opt = 'inv_gauss’ is specified to introduce variability in recruitment. For demonstration purposes, we will set up the following projection scenarios, which include:

  1. Using F40F_{40} for projections, where an HCR is applied to adjust the F40F_{40} in each projection year,
  2. Using F=0F = 0 for projections.

Let us first redefine our single region inputs to use in our projection function, but also define the number of simulations to conduct (1000):


data("sgl_rg_sable_rep") # read in single region report
data("sgl_rg_sable_data") # read in single region data 

# Setup necessary inputs
n_sims <- 1e3 # number of simulations to conduct
t_spawn <- 0 # spawn timing
n_proj_yrs <- 15 # number of projection years
n_regions <- 1 # number of regions
n_ages <- length(sgl_rg_sable_data$ages) # number of ages
n_sexes <- sgl_rg_sable_data$n_sexes # number of sexes
n_fish_fleets <- sgl_rg_sable_data$n_fish_fleets # number of fishery fleets
do_recruits_move <- 0 # recruits don't move
terminal_NAA <- array(sgl_rg_sable_rep$NAA[,length(sgl_rg_sable_data$years),,], dim = c(n_regions, n_ages, n_sexes)) # terminal numbers at age
terminal_NAA0 <- array(sgl_rg_sable_rep$NAA0[,length(sgl_rg_sable_data$years),,], dim = c(n_regions, n_ages, n_sexes)) # terminal numbers at age
WAA <- array(rep(sgl_rg_sable_data$WAA[,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # weight at age
WAA_fish <- array(rep(sgl_rg_sable_data$WAA[,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes, n_fish_fleets)) # weight at age for the fishery
MatAA <- array(rep(sgl_rg_sable_data$MatAA[,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # maturity at age
fish_sel <- array(rep(sgl_rg_sable_rep$fish_sel[,length(sgl_rg_sable_data$years),,,], each = n_proj_yrs), dim = c(n_regions, n_proj_yrs, n_ages, n_sexes, n_fish_fleets)) # selectivity
Movement <- array(rep(sgl_rg_sable_rep$Movement[,,length(sgl_rg_sable_data$years),,], each = n_proj_yrs), dim = c(n_regions, n_regions, n_proj_yrs, n_ages, n_sexes)) # movement
terminal_F <- array(sgl_rg_sable_rep$Fmort[,length(sgl_rg_sable_data$years),], dim = c(n_regions, n_fish_fleets)) # terminal F
natmort <- array(sgl_rg_sable_rep$natmort[,length(sgl_rg_sable_data$years),,], dim = c(n_regions, n_proj_yrs, n_ages, n_sexes)) # natural mortality
recruitment <- array(sgl_rg_sable_rep$Rec[,20:(length(sgl_rg_sable_data$years) - 2)], dim = c(n_regions, length(20:length(sgl_rg_sable_data$years) - 2))) # recruitment values to use for mean recruitment calculations or inverse gaussian parameterization
sexratio <- array(0.5, dim = c(n_regions, n_proj_yrs, n_sexes)) # recruitment sex ratio 

We can then define our two projection scenarios to use in this example, as well as arrays to store our projection results in:

# Define the F used for each scenario 
proj_inputs <- list(
  # Scenario 1 - Using HCR to adjust f40
  list(f_ref_pt = array(sgl_ref_pt$f_ref_pt, dim = c(n_regions, n_proj_yrs)),
       b_ref_pt = array(sgl_ref_pt$b_ref_pt, dim = c(n_regions, n_proj_yrs)),
       fmort_opt = 'HCR'
  ),
  # Scenario 2 - F is set at 0
  list(f_ref_pt = array(0, dim = c(n_regions, n_proj_yrs)),
       b_ref_pt = NULL,
       fmort_opt = 'Input'
  )
)

# store outputs
all_scenarios_f <- array(0, dim = c(n_regions, n_proj_yrs, n_sims, length(proj_inputs)))
all_scenarios_ssb <- array(0, dim = c(n_regions, n_proj_yrs, n_sims, length(proj_inputs)))
all_scenarios_catch <- array(0, dim = c(n_regions, n_proj_yrs, n_fish_fleets, n_sims, length(proj_inputs)))

Next, we can then run our projections by iteration through our scenarios, as well as the number of simulations to conduct. Note that we switch from fmort_opt = 'HCR' to fmort_opt = 'Input' when going from scenario 1 to scenario 2, given that scenario 2 does not use an HCR (the HCR used is the threshold control rule defined earlier).

set.seed(123)
for (i in seq_along(proj_inputs)) {
  for (sim in 1:n_sims) {

    # do population projection
    out <- Do_Population_Projection(n_proj_yrs = n_proj_yrs, # number of projection years
                                    n_regions = n_regions, # number of regions
                                    n_ages = n_ages, # number of ages
                                    n_sexes = n_sexes, # number of sexes
                                    sexratio = sexratio, # sex ratio
                                    n_fish_fleets = n_fish_fleets, # number of fleets
                                    do_recruits_move = do_recruits_move, # whether recruits move
                                    recruitment = recruitment, # recruitment values to use to parameterize inverse gaussian
                                    terminal_NAA = terminal_NAA, # terminal numbers at age
                                    terminal_NAA0 = terminal_NAA0, # terminal unfished numbers at age
                                    terminal_F = terminal_F, # terminal fishing mortality at age
                                    natmort = natmort, # natural mortality 
                                    WAA = WAA, # weight at age for spawning
                                    WAA_fish = WAA_fish, # weight at age for fishery
                                    MatAA = MatAA, # maturity at age
                                    fish_sel = fish_sel, # fishery selectivity
                                    Movement = Movement, # movement
                                    f_ref_pt = proj_inputs[[i]]$f_ref_pt, # fishing mortality to use for projection
                                    b_ref_pt = proj_inputs[[i]]$b_ref_pt, # biological reference point to use in HCR
                                    HCR_function = HCR_function, # harvest control rule function
                                    recruitment_opt = "inv_gauss", # stochastic simulation for recruitment
                                    fmort_opt = proj_inputs[[i]]$fmort_opt, # fishing mortality option (either HCR or a user input)
                                    t_spawn = t_spawn # spawn timing
    )

    # store results
    all_scenarios_ssb[,,sim,i] <- out$proj_SSB
    all_scenarios_catch[,,,sim,i] <- out$proj_Catch
    all_scenarios_f[,,sim,i] <- out$proj_F[,-(n_proj_yrs+1)] # remove last year, since it's not used
    
  } # end sim loop
  print(i)
} # end i loop

Finally, we can plot these stochastic simulations to inspect results:

# Get historical SSB
historical <- reshape2::melt(array(rep(sgl_rg_sable_rep$SSB, n_sims),
                                   dim = c(n_regions, length(sgl_rg_sable_data$years), n_sims))) %>%
  mutate(Year = Var2 + 1959,
         Scenario = "FABC (F40)",  # or change to match the scenarios you're plotting
         Type = "Historical") %>%
  rename(Region = Var1, Simulation = Var3, SSB = value)

# Get all scenario projections
scenarios <- reshape2::melt(all_scenarios_ssb) %>%
  mutate(Year = Var2 + 2023,
         Scenario = case_when(
           Var4 == 1 ~ "S1: FABC (F40)",
           Var4 == 2 ~ "S2: F = 0"
         ),
         Type = "Projection") %>%
  rename(Region = Var1, Simulation = Var3, SSB = value)

# expand historical SSB for plotting
scenarios_unique <- unique(scenarios$Scenario)
historical_expanded <- historical[rep(1:nrow(historical), times = length(scenarios_unique)), ]
historical_expanded$Scenario <- rep(scenarios_unique, each = nrow(historical))

# combine
combined_ssb <- bind_rows(historical_expanded, scenarios)

# Plot
combined_ssb %>%
  ggplot(aes(x = Year, y = SSB, group = interaction(Scenario, Simulation), color = Type)) +
  geom_line(alpha = 0.05) +
  facet_wrap(~Scenario, scales = 'free') +
  geom_hline(yintercept = sgl_ref_pt$b_ref_pt, lty = 2) + # b40
  geom_vline(xintercept = 2024, lty = 2) + # projection start
  scale_color_manual(values = c("Historical" = "black", "Projection" = "blue")) +
  theme_bw(base_size = 15) +
  theme(legend.position = 'none')

# Get historical catch
historical <- reshape2::melt(array(rep(sgl_rg_sable_data$ObsCatch, n_sims),
                                   dim = c(n_regions, length(sgl_rg_sable_data$years), sgl_rg_sable_data$n_fish_fleets, n_sims))) %>%
  mutate(Year = Var2 + 1959,
         Scenario = "FABC (F40)",  # or change to match the scenarios you're plotting
         Type = "Historical") %>%
  rename(Region = Var1, Simulation = Var4, Fleet = Var3, Catch = value) %>%
  select(-Var2)

historical$Catch[is.na(historical$Catch)] <- 0

# Get all scenario projections
scenarios <- reshape2::melt(all_scenarios_catch) %>%
  mutate(Year = Var2 + 2023,
         Scenario = case_when(
           Var5 == 1 ~ "S1: FABC (F40)",
           Var5 == 2 ~ "S2: F = 0"
         ),
         Type = "Projection") %>%
  rename(Region = Var1, Simulation = Var4, Catch = value, Fleet = Var3) %>%
  select(-c(Var2, Var5))

# expand historical SSB for plotting
scenarios_unique <- unique(scenarios$Scenario)
historical_expanded <- historical[rep(1:nrow(historical), times = length(scenarios_unique)), ]
historical_expanded$Scenario <- rep(scenarios_unique, each = nrow(historical))

# combine
combined_cat <- bind_rows(historical_expanded, scenarios)

# Plot
combined_cat %>%
  group_by(Year, Scenario, Simulation, Type, Region) %>%
  summarize(Catch = sum(Catch)) %>%
  ggplot(aes(x = Year, y = Catch, group = interaction(Scenario, Simulation), color = Type)) +
  geom_line(alpha = 0.05) +
  facet_wrap(~Scenario) +
  coord_cartesian(ylim = c(0, NA)) +
  scale_color_manual(values = c("Historical" = "black", "Projection" = "blue")) +
  theme_bw() +
  theme(legend.position = 'none')

References

Kapur, M.S., Siple, M.C., Olmos, M., Privitera-Johnson, K.M., Adams, G., Best, J., Castillo-Jordán, C., Cronin-Fine, L., Havron, A.M., Lee, Q., Methot, R.D., Punt, A.E., 2021. Equilibrium reference point calculations for the next generation of spatial assessments. Fisheries Research 244, 106132. https://doi.org/10.1016/j.fishres.2021.106132